Wilhelm Johannsen

Wilhelm Ludvig Johannsen (February 3, 1857 - November 11, 1927) was a Danish botanist. Born in Copenhagen, he worked at the Institute of Agriculture there, before becoming professor of agriculture at the University of Copenhagen. In 1909, he coined the word gene (using the Greek for "to give birth to").

The terms "phenotype" and "genotype" were created by Wilhelm Johannsen in 1909.


 * Reference of Johannsen's major work: Johannsen WL (1909) Elemente der exakten Erblichkeitslehre. Gustav Fischer, Jena
 * Johannsen W (1911) The genotype conception of heredity. Am Nat 45:129-159

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