Lepidodendron

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Lepidodendron
Fossil range: Carboniferous
Scientific classification
Kingdom: Plantae
Division: Lycopodiophyta
Class: Isoetopsida
Order: Lepidodendrales
Family: Lepidodendraceae
Genus: Lepidodendron
Species

See text.

Lepidodendron (also known as the "Scale tree") is an extinct genus of primitive, vascular, arborescent (tree-like) plant related to the Lycopsids (club mosses). They are called Giant club mosses, since they sometimes reached heights of over 30 m, and the trunks were often over 1 m in diameter. They thrived during the Carboniferous period.

Lepidodendron had tall, thick trunks that rarely branched and were topped with a crown of bifurcating branches bearing clusters of leaves. These leaves were long and narrow, similar to large blades of grass, and were spirally-arranged. The vascular system was a siphonostele with exarch xylem maturation.

The closely packed diamond-shaped leaf scars left on the trunk and stems as the plant grew provide some of the most interesting and common fossils in Carboniferous shales and accompanying coal deposits. These fossils look much like tire tracks or alligator skin.

Image:Lepidodendron aculeatum2.jpg
Fossil trunk of Lepidodendron aculeatum showing leaf scars

The scars, or leaf cushions, were composed of green photosynthetic tissue, evidenced by the cuticle covering and being dotted with stomata, microscopic pores through which carbon dioxide from the air diffuses into plants. Likewise, the trunks of Lepidodendrons would have been green, unlike modern trees which have scaly, non-photosynthetic brown or gray bark.

Lepidodendron has been likened to a giant herb. The trunks produced very little, if any, wood. Most structural support came from a thick, bark-like region. This region remained around the trunk as a rigid layer that did not flake off like that of most modern trees. As the tree grew, the leaf cushions expanded to accommodate the increasing width of the trunk.

The branches of this plant ended in cone-like structures. Lepidodendron did not produce seeds like many modern plants. Instead, it reproduced by means of spores. It is estimated that these plants grew rapidly and lived 10-15 years. Some species were probably monocarpic, meaning they reproduced only once toward the end of their life cycle.

Lepidodendron likely lived in the wettest parts of the coal swamps that existed during the Carboniferous period. They grew in dense stands, likely having as many as 1000 to 2000 giant clubmosses per hectare. This would have been possible because they did not branch until fully grown, and would have spent much of their lives as unbranched poles. In its juvenile stages, the trunk was supported by grass-like leaves that grew straight out of the trunk.

By the Mesozoic era, the giant clubmosses had died out and were replaced by smaller clubmosses, probably due to competition from the emerging woody gymnosperms and other plants. Lepidodendron is one of the more common plant fossils found in Pennsylvanian (Carboniferous) age rocks. They are closely related to other extinct genera, Sigillaria and Lepidendropsis.

Trivia

In the 19th Century, due to the reptilian look of the diamond-shaped leaf scar pattern, petrified trunks of Lepidodendron were exhibited at fairgrounds as giant fossil lizards or snakes.

The name Lepidodendron comes from the Greek lepido, scale, and dendron, tree.

See also

References

it:Lepidodendron nl:Lepidodendron ja:リンボク

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Acknowledgement and Attribution Regarding Sources of Content

Some of the initial content on this page may be incorporated in part from copyleft sources in the public domain including wikis such as Wikipedia and AskDrWiki. Drug information for patients came from the The National Library of Medicine. Infectious disease information may have come from the Centers for Disease Control (CDC). Differential Diagnoses are drawn from clinicians as well as an amalgamation of 3 sources: 1.The Disease Database; 2. Kahan, Scott, Smith, Ellen G. In A Page: Signs and Symptoms. Malden, Massachusetts: Blackwell Publishing, 2004:3; 3. Sailer, Christian, Wasner, Susanne. Differential Diagnosis Pocket. Hermosa Beach, CA: Borm Bruckmeir Publishing LLC, 2002:7 .

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